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Noss has awakened. The planet has tasked the wayfarers with recovering long-lost memories from before the Downfall. To complete this quest, they will need to open all sixteen domes. As the party splits, seafaring battles leave Lago-Sterjall stranded in a desert. Deceit breeds beneath the blue sands, and grave dangers threaten to take away all that the wolf holds dear, including his newfound lover, the caracal prince. The dome of the marsupial clade is now within their reach, but even more realms await their arrival-domains where the limits of Sterjall's moral convictions will be tested. Journeying toward the opposite end of the continent, Jiara-Kitjári recruits Banook and his bears to infiltrate the domes ransacked by the Negian Empire. While traversing frozen mountains and forsaken ruins filled with riches, Kitjári's heart makes room for a more valuable treasure, but the woman she loves does not seem to share the same sentiment. All the while, the Red Stag's army advances, now threatening to enslave the behemoths of the Archstone Dome. If the warmongering elk manages to rein in the giants, the Negian Empire will be ready to carry out its final strike.